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This category is a mixed bag of settings, covering both entries that adjust delay times (c2 through c4) and how the shutter release and AE-L buttons interact (c1). I think the latter setting should have been placed in the purple f-coded Controls section. Go figure.
This is another of Nikon’s easily confusing options for controlling how and when autofocus and exposure are activated and locked. The intent is to allow you to separate autofocus and autoexposure activation and locking. All you need to know is: