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There are use cases that might require the ability to translate or proxy between IPv4 and IPv6. For example, translation in the data center is necessary to enable communication between IPv6-enabled hosts in the campus/branch and legacy IPv4-only servers in the data center access layer.
An intermediate device or node (for example, router, firewall, or load balancer) can translate from IPv4 to IPv6 and vice versa. Or the operating system can perform translation at each endpoint.